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Each ionizable group of an amino acid can exist in one of two states, charged or neutral. The electric charge on the functional group is determined by the relationship between its p Ka and the pH of the solution. This relationship is described by the Henderson-Hasselbalch equation. (a) Histidine has three ionizable functional groups. Write the equilibrium equations for its three ionizations and assign the proper p K2 for each ionization. Draw the structure of histidine in each ionization state.
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